You really should not edit any of the .css templates *except* the additional_css.css template. You should add any changes your want to the CSS into that template.
But anyhow, so, editing *any* template at all gives that error? Can you check your /includes/class_core.php file and make sure it is complete and is for 4.2.5? Line 5559 is just a comment line and shouldn't be giving any eval errors.
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