When you disabled the plugins, did you also remove any extra files you uploaded to your server for your modifications? And, you say you clicked through all your styles, but do you have a completely default vbulletin style that you also looked at the site in?
Your style is staying that way for IE because you probably clicked on the style from your admin panel when you were in IE and the cookie got set. Try clicking on the style you want in your admin panel to get your style back in IE.
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