What's posted only removes the underscore from the display, not the actual thread title e.g. (I noticed you're using vBSEO)
Thread title: Hello_World
Thread title would display as: Hello World
Thread URL: http://www.someforum.com/introductions/1684/hello_world.html
to get it to remove from the URL and the display and thread title (for newly posted threads) import the attached plugin.
*Note* I'm not sure if it works for 3.5.x but it should the only difference between the 3.5 plugin and 3.6 plugin is the 3.5 plugin has executionorder="5" removed....
Iain
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